Mechanisms and Strategies for Phytoremediation of Cadmium

INTRODUCTION
    Phytoremediation
    Advantages of  Phytoremediation
    Limitations of Phytoremediation

BIOAVAILABILITY OF CADMIUM 
    Cation Exchange Capacity (CEC)
    pH
    Soil Amendments
    Competitive Cations
    Fertilizer
    Mycorrhizae
    Chelation
       Phytochelatins (PCs)
       Phytochelatin Effectiveness
       Role of Sulfur in PCs
       Oxidative Stress
       Translocation
       Metallothioneins
       Organic Acids
       EDTA / EGTA

CADMIUM TOLERANCE AND
ACCUMULATION IN PLANTS
    Cell Wall Binding
    Reduced Transport
    Compartmentalization
    Chelation
    Phytoextraction factors
       Table 1.  Plant Accumulation
       Hyperaccumulators

CONCLUSIONS

LINKS

BIBLIOGRAPHY

 
Cation Exchange Capacity

     The most important factor regulating Cd bioavailability is the cation exchange capacity (CEC) of the soil. Clay particles, called micelles, are negatively charged and reversably bind (adsorb) positively charged particles (cations) to their surface. Cations such as Cd may be exchanged for H+ on the micelle surface. Cations adsorbed to micelles are not available for plant uptake or groundwater-promoted migration. Conversely, cations not adsorbed to the micelle are available for uptake (Moore, et al. 1995). Different metal cations adsorb to micelles with different affinity, and adsorption among clay soils varies by as much as 21% (Atanassova, 1999). Copper (Cu) binds more tightly than zinc (Zn) which binds more tightly than nickel (Ni) or Cd (Atanassova, 1999). Because of this, the mineral composition of the soil may increase or decrease depending on the presence of other metals. For example, a soil high in Cu would have more bioavailable Cd since Cu inhibits Cd-adsorption to micelles (Atanassova, 1999). 
 
     Furthermore, cation exchange capacity varies among soil types, regulated by the relative amount of clay present. Since only clay binds cations, soils high in sand or silt have lower CEC. Sandy soils have greater cation availability than other soils, but cations are often leached out by water quicker than they can be utilized by plants. For this reason, clay is often added to soils to prevent cation mobilization and/or uptake (Vangronsveld, 1990). In order for phytoextraction of Cd to proceed efficiently, the CEC of the soil must be low enough that significant amounts of Cd are bioavailable, yet high enough that Cd is not so bioavailable that it leads to toxic reactions in the plant. Several strategies for altering Cd bioavailability via manipulation of soil CEC are available.

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Page Created 4-18-00
Sam Cox
Department of Horticulture
Colorado State University
samcox@lamar.colostate.edu
http://lamar.colostate.edu/~samcox/index.htm